In src/keyfile.c:

> @@ -235,6 +235,8 @@ static void init_pref_groups(void)
>  		"use_gio_unsafe_file_saving", TRUE);
>  	stash_group_add_boolean(group, &file_prefs.keep_edit_history_on_reload,
>  		"keep_edit_history_on_reload_125", FALSE);

This needs to be renamed back to keep_edit_history_on_reload without the _125 suffix to reset the default value.

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